P-21

Molecular breeding to improve grain yield in high quality two-row malting barley.
S.E. ULLRICH (1), A. Kleinhofs (1), D. Schmierer (1), D.A. Kudrna (1), V.A. Jitkov (1), and B.L. Jones (2). (1) Washington State University, Pullman, and (2) USDA-ARS, Madison, WI.

Production of high-quality malting barley is limited by competition from higher-yielding feed barley cultivars in the Northwest USA. This persistent problem has not been overcome by conventional barley breeding. The availability of high-quality malting barley cultivars that are competitive in yield with the best feed barley cultivars would benefit barley growers and the malting and brewing industries. The objectives of this study were to use molecular breeding technology to improve two-row malting barley and to verify specific genetic effects. Introgression of two putative yield quantitative trait loci (QTLs) on chromosome 2HL and one on chromosome 3HL from 'Baronesse' into 'Harrington' was attempted via molecular marker assisted backcrossing (BC). The intended result is the development of lines with yield equal to or greater than Baronesse and malting quality equal to or greater than Harrington; that is lines with Baronesse yield QTLs in a Harrington genetic background. Sixty-four of 112 or 57 percent of the BC lines equaled or bettered the yield of Baronesse in the 2000 field tests. Over three locations in the 2001 tests, 17 of 28 or 61 percent of the BC lines advanced equaled the yield of Baronesse. Considering all tests the highest-yielding BC lines averaged 97 to 105 percent of Baronesse. All but one of the highest-yielding BC lines have at least one of the three yield QTLs from Baronesse. Twenty-one of 28 or 75 percent of the BC lines advanced to the 2001 field trials had malt quality scores higher than Harrington. The five lines advanced to 2002 state yield trials had malt extracts ranging from 80.7 to 81.4 percent vs 80.7 and 78.1 percent for Harrington and Baronesse, respectively. The high proportion of high-yielding and high-quality lines is expected in molecular breeding, whereas it is not in conventional breeding. Genotypic selection is expected to be more precise than phenotypic selection. Extent of the Harrington genetic background in the BC lines partially explains variation among the lines. The interaction or epistasis among the genes of both quantitative characters, yield and malting quality, also play a role in the variation observed. Tests thus far are very encouraging for developing a breakthrough two-row malting cultivar with high yield and high quality by molecular breeding.

Steve Ullrich is a Professor in the Dept. of Crop and Soil Sciences at Washington State University. He came to WSU in 1978 after obtaining the Ph.D. degree from Montana State University. Current research includes molecular genetics study of barley adaptation and quality traits and cultivar development. He also teaches and advises undergraduate and graduate students. He has been active in the North American Barley Genome Mapping Project, formally as a Steering Committee member and currently as a researcher. He is a Northwest representative of the National Barley Improvement Committee and the NBIC/USDA-ARS Barley Crop Germplasm Committee.

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